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Penis fencing is a mating behavior engaged in by many species of flatworm, such as Pseudobiceros hancockanus.

Of these diseases, hookworm, Stoll’s “great infection of mankind,” is one of the most debilitating and widely disseminated.

For example, the concept of mammals (such as lions and certain species of seal) who have "harems" has been commonly reinterpreted as females tolerating a single male simply due to access to resources his leadership provides, while having more than one is simply bothersome to the group after a certain age.

However, this doesn't mean fewer males contribute to the species; many live on their own or in bachelor groups, taking what they can get.

Other species will alternate which partner transfers sperm, engaging in multiple bouts of fencing with the same partner over time. modesta, bilateral sperm transfer is the most common, especially in similarly-sized mate pairs.

The term also applies to homosexual activity between two males among bonobos; same-sex genital-genital rubbing is used in bonobo society to cement bonds, reduce conflict, and express communal excitement over food.

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Pairs can either compete, with only one individual transferring sperm to the other, or the pair can transfer sperm bilaterally.

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